Critically endangered freshwater pearl mussels have been discovered living in two Scottish lochs.
Scientists said the protected species, which is vulnerable to pollution and poaching, had previously only been found in rivers.
Researchers using remotely-operated underwater drones and snorkelling equipment made the discoveries in lochs in Sutherland and the Trossachs.
Scotland’s nature agency, NatureScot, said they likely entered the lochs by clinging to the gills of fish.
Early in their life cycle the mussels live harmlessly on young salmon and trout, before dropping off to live on riverbeds.
NatureScot’s Iain Sime said other lochs would now be surveyed for the species.
He said: “Other mussel species, including the much more widespread swan and duck mussels, are known to live and breed in Scottish lochs, but up until now we’ve had no evidence to suggest that pearl mussels routinely do this too.
“This project was an exciting first step in exploring our lochs and we’re keen to do further surveys to better understand more about this critically endangered species.”
He added: “We don’t yet know whether they can breed in lochs and we want to learn more about a loch pearl mussel’s relationship with their host fish.”
Freshwater pearl mussels are similar in shape to common marine mussels, but can grow larger and live for up to 130 years in fast-flowing rivers.
In history, Julius Caesar’s admiration of pearl mussels is cited by his biographer as a motive for the first Roman invasion of Britain in 55BC.
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